A variety of molds will grow on the combs when the moisture in the hive gets too high. The moisture gets too high when there are not enough bees to fan it away. I suppose the confusion arises because these moldy combs frequently contain dead bees that are also covered with mold, so it looks like the mold killed everything.
According to various sources, one of the molds frequently found on combs is Penicillium waksmanii which can actually inhibit the growth of certain bacteria, including American foulbrood. Other molds are usually present as well, as evidenced by the different colors—usually blue, white, yellow, or gray.
If you have a dead hive loaded with mold you will be tempted to discard the comb, but it is usually not necessary. The first thing you need to do is decide how the colony died or became weak. If you can eliminate American foulbrood and Nosema ceranae, then you can re-use the combs.
https://www.honeybeesuite.com/what-to-do-with-moldy-combs/
Physics for beekeepers: mold in a beehive https://www.honeybeesuite.com/physics-for-beekeepers-mold-in-a-beehive/
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