According to Acinq 'Eclair' functions like any light-client wallet and additionally, when Segwit gets activated the wallet will utilize the protocol for further LN functionality
Acinq’s Eclair wallet separates on-chain transactions from off-chain Lightning Network transactions.
The wallet’s user interface records both on-chain and off-chain transactions with different icons highlighted in the transaction log. Moreover, if users obtain a QR code for payment, the software will detect whether it is a regular bitcoin address or an LN payment request. There’s no need to reset the app to accept LN channels as the platform will recognize and filter through both types of sending procedures.
“So basically, you are free to use the wallet on-chain only, but if you wish to pay less fee and make instant payments, just open one or more channels, and you’ll be able to pay through Lightning,” the startup details
Credits:- (Bitcoin.com & Acinq)